The herd embarks on a quest to find Buck, who reunites them after an asteroid strikes their home, threatening the planet. Along the way, they meet Shangri Llama and her immortal animal community, Geotopia, where Sid inadvertently destroys the city and helps save it from an asteroid by using its crystals to deflect it away. With Buck's plan working, the herd returns home, reconciling some relationships, and celebrating with weddings.
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